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- 国際学会発表:The genetic basis of the evolution of self-compatibility in wild Petunia axillaris
- 渡航報告:オランダ・Wageningen University & Research
- 国際学会発表:Population genomics reveals the evolutionary history of climate adaptation in Japanese Arabidopsis halleri
- 渡航報告:イギリス・John Innes Centre
- 渡航報告:オーストリア・オーストリア科学アカデミーGMI(Magnus Nordborg Lab)